Enterosora - Polypodiaceae

Enterosora barbatula (Baker) Parris






Polydium barbatulum Baker
Grammitis barbatula (Baker) Ching
Polypodium poolii Baker
Grammitis poolii (Baker) Copel.
Polypodium uluguruense Reimers
Grammitis uluguruensis (Reimers) Copel.

Common name


Rhizome short-creeping, sometimes branched, dorsiventral, stipes in 2 rows, 0.5–2 mm apart in each row, articulated to rhizome, phyllopodia 0.4–0.9 mm high; rhizome scales narrowly lanceolate in outline, with glandular hairs ± 0.1 mm long on margin. Stipe 3.2–10.8 cm long, with simple eglandular hairs 1.2–2.5 mm long. Lamina narrowly elliptic to linear-elliptic in outline, 7.3–32+ cm long, 0.7–1.8 c wide, rounded to bluntly pointed at apex, cuneate to long-attenuate at base, smaller fronds entire, larger fronds pinnately divided to 3–4 mm from mid-vein, up to 1/2 of lamina width, lobes up to 10 mm long; texture spongiose-coriaceous to coriaceous; with simple eglandular hairs 1–2.1 mm long on both surfaces of lamina and on margin; lateral veins 1–3-forked or pinnately branched in lobes, free or anastomosing, especially near margin in larger fronds, sometimes along mid-vein. Sori in 2–6 rows, 1–3 on each side of mid-vein, in apical 1/5 to ± throughout lamina, 14–106 in first row (nearest mid-vein), 2–20 in second row, 1–7 in third row.




Tree trunks, sometimes under deeply shaded leaning tree bole, or on overhanging wet rocks, in montane forest on basement complex rocks (granulites).

Distribution worldwide

Africa, Madagascar, Mauritius, Réunion.

Distribution in Africa

Tanzania .

Growth form

Epiphytic, lithophytic.


  • Parris, B.S. (2005) Grammitidaceae.Flora of Tropical East Africa, Pages 13 - 15. (Includes a picture).
  • Roux, J.P. (2009) Synopsis of the Lycopodiophyta and Pteridophyta of Africa, Madagascar and neighbouring islands. Strelitzia 23, South African National Biodiversity Institute, Pretoria. Page 153.